Locule: A chamber within an ovary, anther, sporangium, or fruit; in ovary and fruit, usually corresponding to a carpel.*
In this tomato cross section, the locules are the areas filled with seeds and gel, surrounded by the tomato’s flesh.
*Pell, S.K., Angell, B. (2016). A botanist’s vocabulary: 1300 terms explained and illustrated. Portland: Timber Press.